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Gary Karpen is a Professor specializing in the fields of Cell Biology and Developmental Physiology at the University of California, Berkeley. He leads research focused on understanding the mechanisms of chromosome structure, function, and inheritance, with a particular emphasis on the role of chromatin in gene regulation and cellular organization. Karpen's contributions include major insights into the dynamics of chromosomal behavior during cell division and the ways in which chromosomal abnormalities can lead to various diseases. He is committed to both advancing scientific knowledge and mentoring a new generation of scientists in the field. His active lab engages in interdisciplinary approaches, collaborating with both national and international institutions to explore the complexities of cellular processes. Throughout his career, Karpen has been recognized for his research excellence and dedication to teaching.
The Mathematics Subject GRE is required for the Fall 2026 admissions cycle. General GRE is optional.