Dr. Mark Goulian

Professor

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Biography

Mark Goulian is the Charles William L. Day Distinguished Professor in the Department of Biology at the University of Pennsylvania. He received his Ph.D. from Harvard University in 1990. His research focuses on the mechanisms that bacteria employ to adapt to changes in their environment, particularly biochemical circuits that enable Escherichia coli and related bacteria to sense and respond to intra- and extracellular cues. Goulian is especially interested in how these circuits function, evolve, and interact with pathways within the cell. His current research interests include the adaptation of bacteria to changes in the availability of oxygen and nutrients, the role of bacterial phosphohistidine phosphatases, and the processes that underpin commensal colonization in animal hosts.

Research Interests

Requirements for University of Pennsylvania

Doctorate Program
Requirements
GPA Requirement
Required:3.6
GRE General
Verbal
Required:162
Quantitative
Required:162
Overall
Required:162
GMAT
Total Score
Required:728
Overall
Required:728
TOEFL
Total
Required:115
Prerequisites
Bachelor's degree or equivalent Strong quantitative background
Application Checklist
  • Academic Transcripts
  • Letters of Recommendation (2-3)
  • Resume/CV
  • Statement of Purpose
  • Essays
Specialization Notes

Wharton Doctoral programs cover fields like Finance, Marketing, Management, and Operations, Information and Decisions.