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Stefan Howorka studied Biochemistry at the University of Vienna and conducted his PhD research on bacterial S-layer proteins under the supervision of Professor Werner Lubitz at the Vienna BioCenter and Professor Hagan Bayley at Texas A&M University. After completing his PhD, he underwent postdoctoral training focused on membrane protein nanopores and their rational redesign for biosensing applications, also under Professor Bayley at Texas A&M. He spent three years at an Austrian biotech incubator specializing in single-molecule sensing before joining University College London (UCL) as a lecturer in the Department of Chemistry in 2005. He was promoted to Professor of Chemical Biology in 2016, reflecting his contributions to the field.